[R] MANOVA summary.manova(m) :" residuals have rank"

[Ozgul Inceoglu]

Dear All, I am trying to perform MANOVA. I have table with 504 columns(species) and 36 rows) with two grouping (season and location)
 
Zx <- Z[c(4:504)]
Zxm <- as.matrix(Z)
m<- manova(Zxm~Season*location, data=Z)

when I do summary.aov, I get respond for each species but summary.manova
summary.manova(m) :" residuals have rank" 24<501.

What can it be the reason for this error message?

Thank you,

Ozgul

 Below you can see part of the table.
name	Season 	location	Acetobacter	Aerococcus	Alishewanella	Amaricoccus
xls-nord-01	J	w	0	0,024078979	0	0
bxls-sud-01	J	w	0	0	0	0
brux-nord-04	A	w	0	0	0	0
brux-sud-04	A	w	0	0	0	0
br-nord-07	Ju	w	0	0	0	0
br-sud-07	Ju	w	0	0	0	0
b-nord-10	O	w	0	0	0	0
bsud-10	O	w	0,107836089	0	0,107836089	0,035945363
Z1-01	J	u	0	0	0	0,040567951
Z3-01	J	u	0	0	0	0
Z5-01	J	d	0,023116043	0	0	0
Z7-01	J	d	0,014130281	0	0	0
Z9-01	J	d	0	0	0	0
Z10-01	J	d	0	0	0	0
Z12-01	J	d	0	0	0	0
Z1-04	A	u	0	0	0	0
Z3-04	A	u	0	0	0	0
Z5-04	A	d	0	0	0	0
Z7-04	A	d	0	0	0	0
Z9-04	A	d	0	0,013839873	0	0
Z10-04	A	d	0	0	0	0
Z12-04	A	d	0	0	0	0
Z1-07	Ju	u	0	0	0	0
Z3-07	Ju	u	0	0	0	0
Z5-07	Ju	d	0	0	0	0
Z7-07	Ju	d	0	0	0	0
Z9-07	Ju	d	0	0	0	0
Z10-07	Ju	d	0	0	0	0
Z12-07	Ju	d	0	0,022301517	0	0
Z1-10	O	u	0	0	0	0
Z3-10	O	u	0	0	0	0
Z5-10	O	d	0	0	0	0
Z7-10	O	d	0	0	0,052924054	0
Z9-10	O	d	0	0	0,035050824	0
Z10-10	O	d	0	0	0	0,040783034
Z12-10	O	d	0	0	0	0